營養階段為植物發育時的主要根基，糖類更為生長代謝重要來源。前期研究中發現 SlSWEET1a 及 SlSWEET1c 糖轉運蛋白分別高度表現在幼葉及根，可能參與植物營養階段的糖分配。為驗証此假說，本研究建立 RNAi 轉殖株進行生長功能觀察及糖含量分析，結果發現在 2-3 週營養生長植株中，SlSWEET1a 表達下降對幼葉及成熟葉的糖累積能力並無顯著影響，然而植物進入老齡 (9-10週) 階段時，表達下降的老葉累積顯著較多糖類，推測其角色可能為植物老化時將糖類運往其它積儲器官。在 SlSWEET1c 的功能分析上，發現SlSWEET1c - GFP表現細胞膜上而SlSWEET1c - GUS則高度表現於根部周鞘或是內皮層，顯示SlSWEET1c可能參與從根部糖份的卸載。進一步建立 RNAi 轉殖株進行糖含量分析，發現SlSWEET1c 表達下降並無影響 2 週大植株根的糖份累積，但在生殖階段可能影響糖類從成熟葉送至根部，且植株種子數量減少。為瞭解析供源及積儲之間的糖平衡關係，本研究中更建立 SlSWEET1a - RNAi 及 SlSWEET1c - RNAi 轉殖株的 T1雜交後代，發現以 SlSWEET1a - RNAi 為父本的雜交植株中發現花粉的受粉率較低，導致種子數量明顯減少，推測 SlSWEET1a 亦可能與花粉糖類供給有關，仍待未來研究。

Vegetative growth determines the productivity of reproductive growth, where sugars are major source of metabolism. Previously, high expression of SlSWEET1a and SlSWEET1c sugar transporters in vegetative leaves and roots, respectively, suggests that these two genes may function in sugar allocation during vegetative growth. To examine the hypothesis, this study established RNAi transgenic tomato plants to examine growth and sugar contents. It was found that reduced expression of SlSWEET1a did not consistently affect sugar contents of young and mature leaves during vegetative ages (2-3 weeks old). Yet, at old stages (9-10 weeks old), reduced expression of SlSWEET1a resulted in sugar accumulation in old leaves, indicating that SlSWEET1a may function to allocate sugars to other sink organs during senescing. In terms of functional assay of SlSWEET1c , localization of GFP in plasma membrane and GUS fusion proteins in pericycle or endodermis cells indicated that SlSWEET1c may participate in sugar unloading to root cells. We further established SlSWEET1c-RNAi transgenic plants to analyze growth phenotypes and sugar contents. Results showed that reduced expression of SlSWEET1c did not affect sugar accumulation during the vegetative stage (2 week old), but affect reproductive growth of seeds. To further understand the sugar homeostasis between source and sink organs, we established F1 seeds of SlSWEET1a-RNAi and SlSWEET1c-RNAi lines. These lines will be investigated in the future.

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